With open cooling in fresh water, much above 160 degrees ƒ actual in the water jacket of the motor will allow bubbles to form on the cylinder/jacket walls. The temperatures go up on the cylinders and heads because of that. Hence you see a lot of 140 and 150ƒ thermostats on open cooled systems. With closed cooling the system maintains a 15 psi pressure and those bubbles won't form below 220ƒ. So those systems can have 190-210 thermostats and the fuel will burn a little more efficiently.
Rick